The Dytiscidae (Say it: die-TISS-uh-dee) – based on the Greek dytikos (δυτικός), "able to dive" – are the diving beetles, a family of water beetles. They occur in virtually any freshwater habitat around the world, but a few species live among leaf litter. The adults of most are between 1 and 2.5 cm (0.4–1.0 in) long, though much variation is seen between species. The European Dytiscus latissimus and Brazilian Megadytes ducalis are the largest, reaching up to 4.5 cm (1.8 in) and 4.75 cm (1.9 in) respectively. In contrast, the smallest is likely the Australian Limbodessus atypicali of subterranean waters, which only is about 0.9 mm (0.035 in) long. Most are dark brown, blackish, or dark olive in color with golden highlights in some subfamilies. They have short, but sharp mandibles. Immediately upon biting, they deliver digestive enzymes. The larvae are commonly known as water tigers. The family includes more than 4,000 described species in numerous genera.