The European jaguar (Panthera gombaszoegensis) lived about 1.5 million years ago, and is the earliest known Panthera species from Europe. Fossil remains were first known from the Olivola site in Italy and under the synonym Panthera toscana from other Italian localities. Later specimens have been found in England, Germany, Spain, France, and the Netherlands. Sometimes it is recognized as a subspecies of Panthera onca, the jaguar. European jaguars were larger than modern-day jaguars found in South America, such as Pantanal and Peruvian jaguars. With a bodyweight between 70 and 210 kg (154 to 463 lbs), they were therefore probably capable of bringing down larger prey. A form similar to Panthera gombaszoegensis has been found dating from early Pleistocene East Africa and had both lion- and tiger-like characteristics.